Journal of Ecology 2013, 101,58–67 doi: -
FO, Northern Ireland, UK; CEH Wallingford, Maclean Building,
Crowmarsh Gifford, Wallingford, Oxon, OX10 8BB, UK; 11Department of Biosciences, College of Science, Swansea
University, Singleton Park, Swansea, SA1 8PP, UK; 12Centre for Ecology and Conservation, College of Life and
Environmental Sciences, University of Exeter, Cornwall Campus, Penryn, Cornwall, TR10 9EZ, UK; 13School of Life
Sciences, University of Sussex, Falmer, Brighton, Sussex, BN1 9QG, UK; 14Institute of Biological and Environmental
Sciences, University of Aberdeen, Aberdeen, AB14 3UU, UK; 15School of Environment, Natural Resources &
Geography, Thoday Building, Bangor University, Bangor, LL57 2UW, UK; 16School of Life Sciences & Maths Institute
17
University of Warwick, Gibbet Hill Road, Coventry, CV4 7AL, UK; Division of Ecology, Evolution and Genetics,
18
Research School of Biology, Australian National University, Canberra, ACT 0100, Australia; School of Biology,
University of Leeds, Leeds, LS1 9JT, UK; 19Environmental Change Institute, School of Geography and the
Environment, University of Oxford, Oxford, OX1 3QY, UK; 20Marine
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